The ground floor is the home of Darwinian Creatures. Such creatures are individually hardwired, without the capacity to change in response to their environments; however, they vary from one another and so the better adapted phenotypes will proliferate in a given environment. So, while individuals may have little or no autonomy, the species (if we may speak of species here) can handle the evolutionary work of autonomy.
Alvaro Moreno and Asier Lasa argue that this evolution depends upon the emergence of internal autonomy or the decoupling of various organic systems from one another.
If these accounts of the evolution of mind are correct, then the capacity of human beings to make autonomous choices, individually or collectively, is a result of the emergence of increasingly sophisticated dimensions of autonomy over the course of evolutionary history. A human person can decide to run before ever she begins to run. She can begin to run before the autonomic process that control her inner nervous system can anticipate or begin to react to her decision. However, to understand human autonomy in its full meaning, we must recognize another decoupling or bifurcation. Just as biological evolution advances by the increasing autonomy of systems internal to the individual organism, so the evolution of social behavior advances by an increasing autonomy between and within social groups. So one cannot fully understand moral autonomy without coming to grips with the tension between the autonomy of the individual and the autonomy of the tribe.